TYPE: Ecuador. Loja: Bajando del puerto de Zamora a Loja, ca. 1 km abajo del puerto, 3°59'08"S 79°05'35"W, 3000 m, 28 January 2004, E. Hágsater & Lester Lapo 14070 (holotype, AMO; xerox copies, AMES, LOJA, MO, QCA, QCNE, SEL, USM; illustration of type).